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Todites coronatus
Todites coronatus Naug. Paleontol. J., 47(12): 1412. Dec 2013
- Name
- Todites coronatus
- Rank
- Species
- Generic Name
- [Genus] Todites
- Authors (Pub.)
- Naugolnykh S. V.
- Publication
- Permian ferns of western Angaraland [2013/12]
- Journal
- Paleontological Journal
- Volume
- 47
- Issue
- 12
- Page number
- 1412
- Year
- 2013
- Fossil Status
- foliage (fertile)
- Stratigraphy
- Roadian
- Strat. comment
- Kazanian Stage
- Location
- Pechora Coal Basin, borehole IK 657, 532.0 m of depth, Komi Republic, Russian Federation
- Paleoregion
- Angarida
Data for Holotypus
- Repository
- Geological Institute, Russian Academy of Sciences, Moscow, Russian Federation
- Repository Number
- 4851/203a
- Diagnosis
- Fern with fertile or partly fertile last
order pinnae with pecopteroid pinnules. Frond rachis
thick, robust, having central adaxial furrow. Sterile
pinnules pecopteroid, connected by their bases and
forming limb (wing), surrounding rachis of last order pinna. Pinnule midveins distinct, extending for three-fourths of pinnule extent and then dichotomizing once
or twice (in basal part of pinna), only once (in middle
part of pinna), or simple (in apical part of pinna). Pinnule apex round or blunted. Fertile pinnules pecopteroid, but sometimes constricted at base. Each fertile pinnule with up to 50 sporangia positioned on pinnule
abaxial surface. Sporangia clavate or fusiform, consisting of two cell types. Basal and middle parts of sporangium consisting of relatively small elongated cells with thin walls. Apical part of sporangium consisting
of larger cells with thick walls. Thick-walled apical cells forming a crownlike structure surrounding stomium of sporangium and responsible for opening stomium. Spores in situ of Osmundacidites type, ranging
from round to subtriangular, up to 50 μm in diameter,
with exine having small spines.
