Global registry of scientific names of fossil organisms covered by the International Code of Nomenclature for Algae, Fungi, and Plants and the International Code of Zoological Nomenclature © 2014-2022

IDNAME species

Knorripteris taylorii

Knorripteris taylorii Galtier, C.J. Harper, Rößler, Kustat., M. Krings in M. Krings, C.J. Harper, Cúneo, G.W. Rothwell Transf. Paleobot. 196. 20 Jul 2018
Knorripteris taylorii
Generic Name
[Genus] Knorripteris
Authors (Pub.)
Galtier J. Harper C. J. Rößler R. Kustatscher E. Krings M.  
Enigmatic, structurally preserved stems from the Triassic of Central Europe: a fern or not a fern? [2018/7]
Authors (Book)
Krings M. Harper C. J. Cúneo N. R. Rothwell G. W.  
Transformative Paleobotany
Page number
Fossil Status
Strat. comment
Precise age unknown. Original label indicates uppermost Muschelkalk or lowermost Keuper
Precise locality unknown. Collected east of the city of Schweinfurt (Schonunger Bucht), rom sands along the left bank of the river Main, Bavaria, Germany
Eurasia (Europe)
Data for Holotypus
Bayerische Staatssammlung für Paläontologie und Geologie, Munich, Germany
Repository Number
Specimen pieces and slides SNSB-BSPG 1968 I 97AeE (specimen pieces) and SNSB-BSPG 1968 I (ex 97) 269e273 (thin sections)
Data for Paratypus
Museum für Naturkunde, Chemnitz, Germany
Repository Number
Specimen and slides under acquisitions K4549AeC, K4549-DS1, and K4549-DS2
Stems up to 60 mm in diameter show acropetal decrease in diameter. Rounded base with central vascularized scar is suggestive of a zone of attachment in one specimen 8.5 mm long. Leaf bases are recognizable as narrow ridges (2 mm wide), helically arranged. Prominent parenchymatous cortex up to 25 mm thick, relatively uniform, has no evidence of zonation. Parenchyma cells are polygonal, often isodiametric, and sometimes showing vertical alignment. Cells vary in diameter from 10 to >200 mm; intercellular spaces exceed 100 mm diameter. Smallest cells (10e20 mm) occur in leaf bases, around the outgoing leaf traces, and in continuous darker zone bounding phloem. Discontinuous phloem cylinder (2 mm thick and 8e16 mm diameter) composed of >10 irregularly shaped strands (each up to 2 mm wide) of thinwalled cells, generally decayed. Phloem parenchyma of small cells (10e20 mm diameter) extend around phloem strands and departing leaf traces, in continuity with innermost cortex. Central circular xylem zone 5e10 mm in diameter, containing numerous flattened strands (up to 2.2 mm wide) of scalariform tracheids surrounded by small-celled parenchyma. Xylem strands, in the continuity of the leaf trace xylem, extend down to center of stem where up to three small strands occur in close contact. There is no evidence of lateral fusion of tracheid bundles, which are enveloped by parenchyma and associated with transfusion tissue. Innermost layers of continuous xylem parenchyma proliferate into cells (20e50 mm diameter and up to 100 mm long) with spiral thickenings (transfusion tissue). Accumulations of this tissue are present only in central “lacunar” xylem zone. Transverse sections of cortex with up to 70 leaf traces are arranged in sets of five, eight, or 13 parastichies. Leaf traces are oval (approximately 1 mm radially  0.5 mm), with the same overall dimensions throughout the cortex. Continuous, multilayered zone of small parenchyma cells encircles and separates tracheid xylem bundles from phloem. Departing leaf traces with xylem bundle are initially flattened, becoming crescent shaped, abaxially curved with abaxial protoxylem. Xylem bundle up to 2 mm wide in phloem zone decreases significantly to <0.5 mm in outer cortex. In contrast, phloem bundles are initially circular (0.5 mm diameter) but slightly increase in size distally. Leaf traces are in the middle cortex frequently with tyloses originating from cells of narrow parenchyma zone between phloem and xylem, and developing inside adjacent tracheid strand. Internodes are very short, approximately 1 mm.

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