Cola amharaensis

Name

Cola amharaensis

Rank

Species

Generic Name

[Genus] Cola

Authors (Pub.)

Pan A. D. Jacobs B. F.  

Publication

The earliest record of the genus Cola (Malvaceae sensu lato: Sterculioideae) from the Late Oligocene (28–27 Ma) of Ethiopia and leaf characteristics within the genus [2009/12]

Journal

Plant Systematics and Evolution

Volume

283

Page number

253

Year

2009

Fossil Status

leaves

Stratigraphy

Chattian

Location

Guang River, Chilga, North Gondar Zone, Amhara Region, north-western Ethiopia; [12° 30′ N, 37° 7′ E]

Paleoregion

Africa (East)

Data for Holotypus

Repository

National Museum of Ethiopia, Addis Ababa, Ethiopia

Repository Number

CH41-P36/CH41-P41

Num

2

Pub.

The earliest record of the genus Cola (Malvaceae sensu lato: Sterculioideae) from the Late Oligocene (28–27 Ma) of Ethiopia and leaf characteristics within the genus [2009/12]

Diagnosis

Entire-margined, mesophyllous leaf with cuneate base, apical pulvinus, and triplinerved basal venation. Primary venation is pinnate, secondary venation is brochidodromous, and tertiary venation is alternate percurrent with admedially ramified veins. Abaxial leaf epidermal cells are rectangular and isodiametric. Stomatal complexes are brachyparacytic and restricted to the abaxial side of the leaf. Anticlinal cell walls are rounded to undulate and have bead-like thickenings. Periclinal cell walls have a stippled texture. Abaxial glandular hairs possess globose or capitate heads and are uniseriate or biseriate and multicellular. Buttressed hair bases are present on the adaxial surface.