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Vladimariales
Order Vladimariales Gordenko Paleontol. J., 44(10): 1282. Dec 2010
- Name
- Vladimariales
- Rank
- Order
- Authors (Pub.)
- Gordenko N. V.
- Publication
- Vladimariales ordo nov. (Gymnospermae) from the Middle Jurassic deposits of the Mikhailovskii rudnik locality (Kursk Region, European Russia) [2010/12]
- Journal
- Paleontological Journal
- Volume
- 44
- Issue
- 10
- Page number
- 1282
- Year
- 2010
- Parent Taxon
- [Class] Ginkgoopsida
- Type
- [Genus] Vladimaria
- Diagnosis
- Plants with peltate, radially sym
metrical seedbearing organs–composite capsules,
formed by fusion of several closed elementary cap
sules, which arranged in whorl on thin axis and
inverted relative to it and base of composite capsule.
Composite capsule dehisced on several lobes along
line of fusion of elementary capsules. Each capsule
contained solitary seed. Seeds, sitting on short stalks,
inverted relative to composite capsule axis. Seeds fusi
form, slightly curved, with chalaza slightly asymmetri
cal and weakly concave in place of stalk attachment;
micropile very long, with apex fused to margins of
small opening in distal margin of capsule. Nucellus
fused with integument more than on two thirds,
megaspore membrane absent. Axis base of composite
capsule sustained by collar on thick stalk, protecting
micropilar tips of seeds. Stalk with several radially
arranged collateral vascular bundles. Their quantity corresponds to quantity of elementary capsules, form
ing composite capsule. Metaxylem tracheids in vascu
lar bundles with scalariform bordered pits. Extensive
resin canals and cavities presented in cortex of stalk,
collar, and composite capsule. Outer epidermis of
composite capsule strongly cutinized, bearing numer
ous perigenous mono or polycyclic stomatal appara
tuses, without sutures, despite of area sustained by col
lar, where radially arranged sutures distinctly
expressed on cuticle. Sutures mark lines of fusion of
capsules. Stalk, bearing collar, upper surface of collar
near its axial part, composite capsule axis, and all area
of composite capsule sustained by collar, papillose;
other surface of composite capsule without papillae.
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